abnormal locomotor behavior, abnormal myelin sheath morphology, abnormal myocardium layer morphology, abnormal nervous system development, abnormal neural plate morphology, abnormal paranode morphology, abnormal response to novel object, abnormal rostral-caudal axis patterning, abnormal rostral-caudal patterning of the somites, abnormal sperm flagellum morphology, abnormal sperm head morphology, abnormal spermiogenesis, abnormal tail development, abnormal vasculogenesis, abnormal visceral yolk sac morphology, abnormal vitelline vasculature morphology, absent somites, behavior/neurological phenotype, cardiovascular system phenotype, decreased embryo size, decreased locomotor activity, decreased myelin sheath thickness, decreased vascular endothelial cell number, embryonic growth arrest, embryonic growth retardation, embryonic lethality between implantation and placentation, complete penetrance, embryonic lethality between somite formation and embryo turning, complete penetrance, embryonic lethality during organogenesis, complete penetrance, embryonic lethality, complete penetrance, incomplete embryo turning, increased grooming behavior, necrospermia, nervous system phenotype, open neural tube, perinatal lethality, complete penetrance, postnatal lethality, incomplete penetrance, premature death, reduced linear vestibular evoked potential, reduced male fertility, short stride length, tonic seizures, wavy neural tube